B. Diversity of Living Things · Diversity of Living ThingsTaxonomy: Ranks, Species and Binomial Nomenclature
B3.1 B2.1 A2.2About 70 minutes
Learning goals- I can explain why biologists classify organisms and why scientific names are more useful than common names.
- I can list the taxonomic ranks from domain to species and use them to compare how closely organisms are related.
- I can write scientific names correctly using binomial nomenclature.
- I can explain the biological species concept and its limits.
Words to know
taxonomy
The science of describing, naming and classifying organisms.
taxon (plural: taxa)
Any named group of organisms at any rank, such as the class Mammalia or the genus Acer.
taxonomic rank
A level in the classification hierarchy: domain, kingdom, phylum, class, order, family, genus, species.
binomial nomenclature
The two-part naming system for species, made standard by Carl Linnaeus: genus name + specific epithet, e.g. Homo sapiens.
genus (plural: genera)
A group of closely related species. The first part of a scientific name.
specific epithet
The second part of a scientific name. On its own it is not a species name.
biological species concept
A species is a group of populations whose members can interbreed in nature and produce fertile offspring.
reproductive isolation
Barriers that prevent members of different species from producing fertile offspring together.
What's in a name?
In Canada, a robin is a large thrush with a brick-red breast that pulls worms out of lawns in spring. In Britain, a robin is a much smaller bird from a different family. In Europe, the animal Canadians call a moose is called an elk, while the North American elk (also called wapiti) is a different deer altogether. Imagine two biologists trying to compare research on "elk"!
To avoid this confusion, every species has one scientific name used by scientists in every language. The American robin is Turdus migratorius. The moose is Alces alces. Today you will learn how these names work and how organisms are arranged into a hierarchy of groups.
Why classify?
There are millions of species. Taxonomy is the science of describing, naming and classifying them. A good classification system:
- gives every species one unique, universal name;
- puts organisms in groups that show relationships (organisms in the same group share an ancestor);
- lets you predict features. If you learn that a new animal is a mammal, you can predict it has hair and feeds its young milk.
A bit of history. The Greek philosopher Aristotle (4th century BCE) sorted animals by features such as whether they had red blood and where they lived. In the 1700s the Swedish botanist Carl Linnaeus built a system of nested groups based on shared physical features, and made the two-part binomial name standard (for plants in 1753, for animals in 1758). We still use his naming system. What has changed is how we decide groups: today they are based on evolutionary relationships, using both morphology and DNA.
The taxonomic ranks
Every species belongs to a series of nested groups, called taxa (singular: taxon). From most general to most specific, the eight main ranks are:
Domain → Kingdom → Phylum → Class → Order → Family → Genus → Species
A memory aid: Dear King Philip Came Over For Good Soup.
Each rank is inside the one above it. As you move down, each group contains fewer kinds of organisms, and the members share more characteristics. (Botanists sometimes use the word division instead of phylum.)
The classification of the grey wolf. Each rank sits inside the one above it. Moving down, the groups get smaller and their members are more closely related. Only the genus and species names are written in italics.| Rank | Human | Grey wolf | Coyote | Canada lynx | North American beaver |
|---|
| Domain | Eukarya | Eukarya | Eukarya | Eukarya | Eukarya |
| Kingdom | Animalia | Animalia | Animalia | Animalia | Animalia |
| Phylum | Chordata | Chordata | Chordata | Chordata | Chordata |
| Class | Mammalia | Mammalia | Mammalia | Mammalia | Mammalia |
| Order | Primates | Carnivora | Carnivora | Carnivora | Rodentia |
| Family | Hominidae | Canidae | Canidae | Felidae | Castoridae |
| Genus | Homo | Canis | Canis | Lynx | Castor |
| Species | Homo sapiens | Canis lupus | Canis latrans | Lynx canadensis | Castor canadensis |
Classification of five mammals found in Canada. The lower the rank two organisms share, the more closely related they are.Worked exampleLet's compare relationships using ranks
Use the table. (a) Which two animals are most closely related? (b) Is the Canada lynx more closely related to the grey wolf or to the beaver? (c) What is the lowest rank shared by all five?
- Find the lowest (most specific) rank each pair shares. The lower the shared rank, the closer the relationship.
- (a) The grey wolf and coyote share genus Canis. No other pair shares a genus, so they are the most closely related.
- (b) Lynx and wolf share order Carnivora. Lynx and beaver share only class Mammalia. Order is lower than class, so the lynx is more closely related to the wolf.
- (c) All five share domain, kingdom, phylum and class, so the lowest shared rank is class Mammalia.
(a) grey wolf and coyote (same genus); (b) the grey wolf (same order); (c) class Mammalia.
Binomial nomenclature: the rules
A species name has two parts:
- The genus name, which always starts with a capital letter.
- The specific epithet, which is always all lower case.
The whole name is written in italics when typed, or underlined when handwritten. After the first full use, the genus can be shortened to its first letter: Acer saccharum, then A. saccharum.
Names are in Latin or Latinized words, and they often describe the organism or honour a person or place. Saccharum means sugar; canadensis means "of Canada"; borealis means northern.
The epithet alone is not a name. canadensis is used for the lynx (Lynx canadensis), the beaver (Castor canadensis), the Canada goose (Branta canadensis) and many plants. Only the pair is unique.
Ranks above genus (such as family Canidae) are capitalized but not italicized.
A scientific name is a genus (capital letter) plus a specific epithet (lower case), both in italics. Three maples share the genus Acer because they are closely related, but each has its own epithet.Worked exampleLet's spot and fix naming errors
Correct each name: (a) Canis Lupus (b) ursus americanus (c) Branta (for the Canada goose) (d) Felidae
- (a) The epithet must be lower case and the name italic: Canis lupus.
- (b) The genus must be capitalized: Ursus americanus (American black bear).
- (c) A genus name alone does not name a species. The Canada goose is Branta canadensis.
- (d) Felidae is a family. Family names are capitalized but not italicized: Felidae.
Canis lupus; Ursus americanus; Branta canadensis; Felidae.
What is a species?
The most widely used definition is the biological species concept: a species is a group of populations whose members can interbreed in nature and produce fertile offspring.
A horse and a donkey can mate, but their offspring, a mule, is almost always sterile. So horses and donkeys are separate species. Things that keep species apart, such as different mating seasons, different courtship behaviours or sterile hybrids, are called reproductive isolation.
The biological species concept has limits:
- It cannot be tested on fossils.
- It does not work for organisms that reproduce asexually, such as bacteria.
- Some separate species occasionally produce fertile hybrids (wolves and coyotes in eastern Canada sometimes interbreed).
So biologists also use morphology and DNA to separate species. You will look at how new species form in the Evolution unit.
Canadian taxonomy today
Taxonomy is an active science. At the University of Guelph, Canadian biologist Paul Hebert proposed DNA barcoding in 2003: identifying species from a short, standard stretch of DNA, much like a grocery barcode. His team founded the Biodiversity Institute of Ontario, and barcoding is now used around the world to identify species and find new ones.
Scientific classification is also not the only knowledge system for living things. First Nations, Métis and Inuit languages contain detailed names for local plants, animals and fungi, often grouped by relationships, behaviour, seasons and uses. Many researchers now work with Indigenous knowledge holders, an approach that Mi'kmaw Elder Albert Marshall calls Two-Eyed Seeing: using the strengths of Indigenous knowledge and Western science together.
Careers: taxonomists, museum curators, molecular biologists and conservation biologists all use classification. Most have a university degree in biology, and many have a master's degree or PhD.
Just for funA funny-but-true fact: the moose is Alces alces, the American bison is Bison bison, and the western lowland gorilla is Gorilla gorilla gorilla. A name that repeats the genus is called a tautonym. Animals are allowed to have them, but the rules for plant names forbid them, so you will never meet a plant with a name like Acer acer.
Hands-on activityClassify your kitchen like Linnaeus
You need: about 20 items from a kitchen cupboard or junk drawer (cans, boxes, utensils, spices; no knives), sticky notes or paper slips, and a large table.
- Lay all the items out. Decide on one feature that splits them into two or three large groups (for example, food vs not food). These are your kingdoms.
- Inside each kingdom, split again using a new feature (for example, canned, boxed, bagged). These are phyla. Keep splitting until each item is alone. Label each level with a sticky note.
- Give each item a two-part binomial name with a genus and a specific epithet in Latin-style words you invent, e.g. Cannus tomatoensis. Items in the same small group must share a genus.
- Write your hierarchy as a table, like the mammal table above.
- Ask a family member to add a new item to your system. Does it fit, or do you need a new group?
What to look for: each level uses a clear, observable feature; every item ends up in exactly one place; items sharing a genus are more similar than items sharing only a kingdom. Discuss: your system is based on your chosen features, which is what early taxonomy was like. Modern taxonomy groups organisms by ancestry, not by convenience.
Your turn
1List the eight taxonomic ranks in order from most general to most specific.
Domain, kingdom, phylum, class, order, family, genus, species.
2Give two reasons scientists use scientific names instead of common names.
Scientific names are unique (one name per species) and universal (the same in every language). Common names vary by place and language, and one common name can refer to different species (e.g. robin, elk).
3Write correctly: the genus Branta, epithet canadensis.
Branta canadensis (italics, capital B, lower-case c).
4Using the table, which is more closely related to humans: the beaver or the lynx? Explain.
Neither is closer by this table. Humans share only class Mammalia with both (humans are Primates; the lynx is Carnivora; the beaver is Rodentia). The table cannot separate them; more evidence is needed. (DNA studies actually place primates closer to rodents than to carnivores, which ranks alone could not tell you.)
5Are horses and donkeys the same species? Explain using the biological species concept.
No. They can produce offspring (mules), but mules are almost always sterile. Under the biological species concept, members of a species must produce fertile offspring.
6Why can the biological species concept not be applied to bacteria?
Bacteria reproduce asexually (by binary fission), so "interbreeding" does not define their groups. Scientists use morphology, metabolism and DNA instead.
7What is a taxon? Give an example at the family level.
A taxon is any named group at any rank. Family examples: Canidae (dogs, wolves, foxes), Felidae, Hominidae.
More practice: warm-up
1Who made binomial nomenclature standard?
2In Homo sapiens, which word is the genus?
3Which rank is directly above genus?
4What does canadensis mean?
5Which Canadian scientist proposed DNA barcoding?
Paul Hebert (University of Guelph).
More practice: on your own
1Two organisms share the same order but not the same family. Do they share the same class? The same genus? Explain.
They share the same class (every order is inside one class), but they cannot share a genus (every genus is inside one family, and they are in different families).
2The red fox is Vulpes vulpes and the Arctic fox is Vulpes lagopus. What can you conclude about their relationship? Name their family.
They share the genus Vulpes, so they are closely related species. Like wolves, foxes belong to the family Canidae.
3Find and fix two errors: "The Eastern Hemlock (tsuga canadensis) belongs to the Pinaceae."
The species name should be italicized with a capital genus: Tsuga canadensis. The family Pinaceae should not be italicized. (Common names are not italicized either: eastern hemlock.)
4Eastern wolves and coyotes in Ontario sometimes interbreed and have fertile hybrids. Why does this create a problem for the biological species concept?
The concept says separate species do not produce fertile offspring together. Fertile wolf-coyote hybrids blur that line, so biologists must also use morphology, DNA and ecology to decide where one species ends and another begins.
5Explain how DNA barcoding could help a fisheries inspector who receives a fillet labelled "red snapper."
A fillet has no fins, colour pattern or shape to identify. A short DNA barcode sequence from the tissue can be compared with a reference library to identify the actual species, revealing mislabelling if the fillet is a cheaper fish.
Challenge
1Linnaeus grouped organisms only by physical similarity. Give an example of how this could produce a misleading group, and explain how modern taxonomy fixes it.
Sample: whales were long thought of as "fish" because they swim and have streamlined bodies and flippers. But whales breathe air with lungs, have hair at some life stage and nurse their young with milk, so they are mammals. Similar looks can come from similar lifestyles, not shared ancestry. Modern taxonomy groups organisms by evolutionary relationships, using many traits plus DNA evidence.
2Species X is in the same family as species Y but a different genus. Species Z is in the same genus as Y. Rank the pairs X-Y, Y-Z and X-Z from most to least closely related.
Y-Z (same genus) is closest. X-Y and X-Z both share only the family, since X is in a different genus from both Y and Z, so they are equally related at this level of detail.
3A biologist finds two populations of salamanders that look identical but never interbreed because one breeds in early spring and the other in early summer. Are they one species or two? Discuss using two species concepts.
By the biological species concept, they are likely two species, because a difference in breeding time is reproductive isolation. By a purely morphological concept they would look like one species. DNA evidence would help decide: if the populations' genes are clearly separated, they are probably distinct, look-alike ("cryptic") species.
Check your understandingQuiz
1. Which list is in order from most general to most specific?
2. Which scientific name is written correctly?
3. The grey wolf and the coyote share genus Canis. Which rank must they also share?
4. A mule is sterile. This shows that horses and donkeys:
5. Why is canadensis not a full species name?
6. Modern taxonomy groups organisms mainly by:
For parents and teachersBig idea (B3.1, B2.1): organisms are classified into nested ranks (domain to species) and named with two-part Latin names. Shared ranks show relatedness. The lesson also introduces the biological species concept and a Canadian scientist (Paul Hebert, DNA barcoding) for A2.2.
How to help: The kitchen-classification activity is a good family task; challenge your teen to justify each split. Quiz them with the mnemonic and ask them to write a few scientific names correctly by hand (underlined).
Common difficulties:
- Capitalizing the epithet (Canis Lupus) or forgetting italics/underlining.
- Thinking more ranks shared means less related. The lower (more specific) the shared rank, the closer the relationship.
- Using the epithet alone as if it were a name.
- Thinking any mating means same species. The offspring must be fertile.
Sources: classifications from standard taxonomic references (ITIS / Canadian Museum of Nature); Linnaeus's Species Plantarum (1753) and Systema Naturae, 10th edition (1758); DNA barcoding proposed by Hebert et al. (2003).